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About This Role
Your Impact at LILA
Lila Sciences is building autonomous systems for scientific discovery, and the AI infrastructure behind that work needs security built in from the start. This role sits on the IT & Security team and owns security engineering for Lila's AI and ML systems, from training and inference pipelines to model registries, MCP servers, and agent runtimes.
As the senior individual contributor for AI security, you will translate the AI security roadmap into delivered controls, drive vulnerability remediation to closure, and reduce risk across the full AI lifecycle: data, models, pipelines, and agentic systems. The work is hands on, with technical depth across cloud security, supply chain security, and AI-specific threat models.
The role is deeply cross-functional. You will partner with AI Platform, AI Research, ML Operations, Science, and AI Safety so security shows up at design time rather than after the fact. Success looks like AI infrastructure that stays continuously compliant, measurable reduction in open risk, and engineering teams that treat security as part of how they build.
What You'll Be Building
- Turn the AI security roadmap into delivered controls with measurable outcomes
- Design and implement security controls across training and inference pipelines, model registries, MCP servers, and agent runtimes
- Keep AI infrastructure continuously compliant against internal standards and applicable frameworks
- Drive vulnerability remediation across AI infrastructure, models, and dependencies to closure with system owners
- Reduce risk across data, models, pipelines, and agentic systems through threat modeling, control design, and adversarial assessment
- Partner with AI Platform, AI Research, ML Operations, Science, and AI Safety so security is embedded at design time
What You'll Need to Succeed
- Strong background in security engineering, cloud security, or DevSecOps, with hands-on experience securing production infrastructure
- Working knowledge of AI/ML systems and the ML lifecycle, plus AI-specific risks such as prompt injection, model and data poisoning, insecure agent tooling, and inference-time threats
- Experience with software supply chain security and dependency and vulnerability management
- Experience implementing data security controls (classification, data flow mapping, access control, encryption) in cloud environments
- Proven ability to partner across engineering teams and drive cross-functional remediation to closure
- Comfort operating with ambiguity in a fast-moving research environment
Bonus Points For
- Hands-on AWS security experience
- Experience with SBOM tooling at scale
- Familiarity with AI governance and assurance frameworks such as NIST AI RMF
- Experience with MAESTRO-style threat modeling for agentic systems
About LILA
Lila Sciences is building Scientific Superintelligence™ to solve humankind's greatest challenges. We believe science is the most inspiring frontier for AI. Rather than hard-coding expert knowledge into tools, LILA builds systems that can learn for themselves.
LILA combines advanced AI models with proprietary AI Science Factory™ instruments into an operating system for science that executes the entire scientific method autonomously, accelerating discovery at unprecedented speed, scale, and impact across medicine, materials, and energy. Learn more at www.lila.ai.
Guided by our core values of truth, trust, curiosity, grit, and velocity, we move with startup speed while tackling problems of historic importance. If this sounds like an environment you'd love to work in, even if you don't meet every qualification listed above, we encourage you to apply.
We're All In
Lila Sciences is committed to equal employment opportunity regardless of race, color, ancestry, religion, sex, national origin, sexual orientation, age, citizenship, marital status, disability, gender identity or Veteran status.
Information you provide during your application process will be handled in accordance with our Candidate Privacy Policy.
A Note to Agencies
Lila Sciences does not accept unsolicited resumes from any source other than candidates. The submission of unsolicited resumes by recruitment or staffing agencies to Lila Sciences or its employees is strictly prohibited unless contacted directly by Lila Science's internal Talent Acquisition team. Any resume submitted by an agency in the absence of a signed agreement will automatically become the property of Lila Sciences, and Lila Sciences will not owe any referral or other fees with respect thereto.
Salary Context
This $252K-$374K range is above the 75th percentile for AI/ML Engineer roles in our dataset (median: $175K across 1356 roles with salary data).
View full AI/ML Engineer salary data →Role Details
About This Role
AI/ML Engineers build and deploy machine learning models in production. They work across the full ML lifecycle: data pipelines, model training, evaluation, and serving infrastructure. The role has evolved significantly over the past two years. Where ML Engineers once spent most of their time on model architecture, the job now tilts heavily toward inference optimization, cost management, and integrating LLM capabilities into existing systems. Companies want engineers who can ship production systems, and the experimenter-only role is fading fast.
Day-to-day, you're writing training pipelines, debugging data quality issues, setting up evaluation frameworks, and figuring out why your model performs differently in staging than it did on your dev set. The best ML engineers are obsessive about reproducibility and measurement. They instrument everything. They know that a model is only as good as the data feeding it and the infrastructure serving it.
Across the 8,943 AI roles we're tracking, AI/ML Engineer positions make up 75% of the market. At Lila Sciences, this role fits into their broader AI and engineering organization.
Demand for AI/ML Engineers has been strong and consistent. Unlike some AI roles that spike with hype cycles, ML engineering is a foundational need. Every company deploying AI models needs people who can keep them running, and the gap between research prototypes and production systems keeps growing.
What the Work Looks Like
A typical week might include: debugging a data pipeline that's silently dropping 3% of training examples, running A/B tests on a new model version, writing documentation for a feature flag system that lets you roll back model deployments, and reviewing a junior engineer's PR for a new evaluation metric. Meetings tend to be cross-functional since ML touches product, engineering, and data teams.
Demand for AI/ML Engineers has been strong and consistent. Unlike some AI roles that spike with hype cycles, ML engineering is a foundational need. Every company deploying AI models needs people who can keep them running, and the gap between research prototypes and production systems keeps growing.
Skills Required
Python and PyTorch dominate the requirements. Most roles expect experience with cloud platforms (AWS, GCP, or Azure) and familiarity with ML frameworks like TensorFlow or JAX. RAG (Retrieval-Augmented Generation) has become a top-3 skill requirement as companies integrate LLMs into their products. Docker and Kubernetes show up in about a third of postings, reflecting the production focus of the role.
Beyond the core stack, employers increasingly want experience with experiment tracking tools (MLflow, Weights & Biases), feature stores, and vector databases. Fine-tuning experience is valuable but less common than you'd think from reading Twitter. Most production LLM work is RAG and prompt engineering, not fine-tuning. If you have both, you're in a strong position.
Companies that are serious about AI/ML hiring tend to post specific infrastructure details in the job description: the frameworks they use, their model serving stack, their data pipeline tools. Vague postings that just say 'ML experience required' without specifics are often companies that haven't figured out what they need yet.
Compensation Benchmarks
AI/ML Engineer roles pay a median of $212,000 based on 11,866 positions with disclosed compensation. Senior-level AI roles across all categories have a median of $228,700. This role's midpoint ($313K) sits 48% above the category median. Disclosed range: $252K to $374K.
Across all AI roles, the market median is $213,380. Top-quartile compensation starts at $264,148. The 90th percentile reaches $319,310. For comparison, the highest-paying categories include AI Safety ($253,100) and Research Engineer ($252,000). By seniority level: Entry: $110,408; Mid: $193,200; Senior: $228,700; Director: $272,250; VP: $235,000.
Lila Sciences AI Hiring
Lila Sciences has 3 open AI roles right now. They're hiring across AI/ML Engineer. Positions span Cambridge, MA, US, Cambridge, MA. Compensation range: $374K - $374K.
Location Context
Across all AI roles, 6% (521 positions) offer remote work, while 8,351 require on-site attendance. Top AI hiring metros: New York (2,878 roles, $221,000 median); San Francisco (2,416 roles, $263,811 median); Los Angeles (1,340 roles, $210,000 median).
Career Path
Common paths into AI/ML Engineer roles include Data Scientist, Software Engineer, Research Engineer.
From here, career progression typically leads toward ML Architect, AI Engineering Manager, Principal ML Engineer.
The fastest path into ML engineering is through software engineering with a self-directed ML education. A CS degree helps, but production engineering skills matter more than academic credentials. Build something that works, deploy it, and measure it. That portfolio project is worth more than a Coursera certificate. For career growth, the fork comes around the senior level: go deep on technical complexity (staff/principal track) or move into managing ML teams.
What to Expect in Interviews
Expect system design questions around ML pipelines: how you'd build a training pipeline for a specific use case, handle data drift, or design A/B testing infrastructure for model deployments. Coding rounds typically involve Python, with emphasis on data manipulation (pandas, numpy) and algorithm implementation. Take-home assignments often ask you to build an end-to-end ML pipeline from raw data to deployed model.
When evaluating opportunities: Companies that are serious about AI/ML hiring tend to post specific infrastructure details in the job description: the frameworks they use, their model serving stack, their data pipeline tools. Vague postings that just say 'ML experience required' without specifics are often companies that haven't figured out what they need yet.
AI Hiring Overview
The AI job market has 8,943 open positions tracked in our dataset. By seniority: 175 entry-level, 4,132 mid-level, 3,268 senior, and 1,368 leadership roles (Director, VP, C-Level). Remote roles make up 6% of the market (521 positions). The remaining 8,351 roles require on-site or hybrid attendance.
The market median for AI roles is $213,380. Top-quartile compensation starts at $264,148. The 90th percentile reaches $319,310. Highest-paying categories: AI Safety ($253,100 median, 57 roles); Research Engineer ($252,000 median, 419 roles); AI Engineering Manager ($251,622 median, 51 roles).
Demand for AI/ML Engineers has been strong and consistent. Unlike some AI roles that spike with hype cycles, ML engineering is a foundational need. Every company deploying AI models needs people who can keep them running, and the gap between research prototypes and production systems keeps growing.
The AI Job Market Today
The AI job market spans 8,943 open positions across 16 role categories. The largest categories by volume: AI/ML Engineer (6,739), AI Product Manager (403), AI Software Engineer (402). These three account for the majority of open positions, though smaller categories often have higher per-role compensation because of specialized skill requirements.
The seniority mix tells a story about where AI teams are in their maturity. Entry-level roles (175) are outnumbered by mid-level (4,132) and senior (3,268) positions, reflecting that most companies are past the 'build a team from scratch' phase and need experienced engineers who can ship production systems. Leadership roles (Director, VP, C-Level) total 1,368 positions, representing the bottleneck between technical execution and organizational strategy.
Remote work availability sits at 6% of all AI roles (521 positions), with 8,351 requiring on-site or hybrid attendance. The remote share has stabilized after the post-pandemic correction. Senior and specialized roles (Research Scientist, ML Architect) are more likely to be remote-eligible than entry-level positions, partly because experienced hires have more negotiating power and partly because these roles require less hands-on mentorship.
AI compensation is structured in clear tiers. The market median sits at $213,380. Top-quartile roles start at $264,148, and the 90th percentile reaches $319,310. These figures include base salary with disclosed compensation. Total compensation (including equity, bonuses, and sign-on) runs 20-40% higher at companies that offer those components.
Category matters for compensation. AI Safety roles lead at $253,100 median, while Prompt Engineer roles sit at $141,440. The spread between highest and lowest-paying categories reflects the premium on specialized technical skills versus broader analytical roles.
The most in-demand skills across all AI postings: Python (1,422 postings), Aws (808 postings), Azure (638 postings), Rag (555 postings), Gcp (457 postings), Pytorch (433 postings), Claude (341 postings), Kubernetes (337 postings). Python dominates, appearing in the vast majority of role descriptions regardless of category. Cloud platform experience (AWS, GCP, Azure) is the second most common requirement. The newer entrants to the top skills list (RAG, vector databases, LLM APIs) reflect the shift from traditional ML toward generative AI applications.
Frequently Asked Questions
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